The story begins with Shuji (シュウジ Shūji), a high school student in a Hokkaidō coastal city, walking up to an observatory and reminiscing about his girlfriend, Chise (ちせ); there he finds her exchange diaries that she purposefully left behind. The ensuing story is narrated by Shuji through flashbacks while reading Chise's diary. Chise, a fellow student in his class, declares her love for Shuji at the beginning of the series. However, Chise is very shy and Shuji is insensitive: neither know how to express their feelings very well, but they do indeed have feelings for each other.
One day, while Shuji is shopping in Sapporo, unknown bombers attack the city in broad daylight. He and his friends run for cover, but notice a fast and small flying object shooting down the enemy bombers. Separated from his friends, Shuji wanders through the wreckage - only to stumble upon Chise; here she has metal wings and weapons - apparently grafted onto her body. She tells him she has become the ultimate weapon, without her knowledge or consent, and that she is seen by the Japan Self-Defense Forces (JSDF) as the last hope for defending Japan from imminent invasion. In the anime, it is not apparent why Chise was chosen to be the ultimate weapon or why the country is under attack. It was not until the OVA episodes were released that an explanation for Chise being chosen was offered: her body has the highest degree of compatibility with the weapon system. (In fact, later in the OVA it is mentioned that Chise had been "over-developed" and was so much of a weapon that she would kill anyone regardless of what side they were on. This fact is punctuated by Chise vaporizing the Special Forces building where she first became the ultimate weapon.)
This story, as suggested by its subtitle "The Last Love Song On This Little Planet", is primarily a love story. It focuses primarily on Shuji's and Chise's reactions to her increasingly powerful destructive abilities, and the relationship between the two.
A number of minor characters who do not necessarily know of Chise's role in the war have sub-plots that concern everyday people in the context of war: a woman whose husband is constantly away from home, a school boy who joins the army to protect his girlfriend, a girl whose civilian boyfriend is killed in a bombing, and others.
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